The EGE
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I feel that Boston has less bus service density roaming the streets compared to places like Chicago or NY. Part of it is the BERy-designed hub-and-spoke structure of the network. But still -- despite the extensive subway infrastructure in NY and Chicago, there is still a thick network of buses laid on top of that, and they run frequently. Also both remnants of disassembled streetcar networks.
I can't really say if Boston of 1940 had a similar density of service provided by streetcars, and lost it, or if this is something more fundamental. The street network seems to make things difficult because only the main roads are really usable by buses and they follow the old intercity / square-to-square routes.
And if Boston did get serious about surface transit (which I think they should, since it's probably the only thing we can reasonably do in the near future), they'd still need more space to store and maintain buses.
F-Line beat me to part of this, but here's my thoughts for completeness's sake:
The problem isn't really BERy's network design - it's excellent for getting a large number of people onto a few trunk lines that serve the most desired destinations. Get better connections to the LMA and the Seaport, and do the eat-ya-peas work to boost Red Line frequencies for Kendall access, and that basic system works pretty well. (And yes - Urban Ring, GLX, BLX-Lynn, and second GL trunk subway were all BERy-originated ideas.)
The problem is twofold: the trunk lines are not running at their full capacities, and the surface routes are pitiful. The heavy rail lines only had four-car trains until the last few decades, but fleet sizes were similar or larger (per line length) so headways were a lot smaller. The Red Line ran 155 cars Harvard-Ashmont, and now runs 220 cars Alewife-Ashmont/Braintree with well over twice the track miles. What are now the B and C lines once ran through to Lechmere with two (and sometimes three) car trains of center-entrance cars (aka crowd swallowers).
While the surface routes haven't actually lost much area since the BERy days. Take a look at this 1943 BERy map; the rapid transit routes have changed, but very little of that surface network is no longer around. And the unified system serves a much larger territory. Route numbered 1-121 are mostly BERy routes (save the 52, 59, 62, 67, 70, 70A, and 76 which are ex-Middlesex & Boston); the 130s, 200s, 350s, and 400s are largely ex-Eastern Mass routes, while the 500s are mostly ex-M&B.
A modern 40-foot bus has more or less the same capacity and average speed as a surface-running streetcar in the first half of the 20th century, so it's not unreasonable to directly compare headways. The frequency of streetcar service was absolutely unbelievable. Major trunk lines like Mass Ave north of Harvard (now the 77) and Blue Hill Ave (now the 28 plus some other routes) had headways under a minute at rush hour. Forget real-time data, there's always a streetcar coming. Even with slow sections and bunching (and real-time headway control is a huge area of active research), buses become a lot more attractive if they come ever 5 minutes rather than every 30.
So yes, the realistic answer is a massive expansion of bus capacity (and some routes eventually converted to BRT, trolleybus, and/or streetcars) which means more bus depots. The long-promised large facilities at Wellington and Arborway would help with that a lot. There's still a lot of industrial land in Boston where building bus depots wouldn't be horribly objected to, and several T-owned parking surface facilities (Riverside, Quincy Adams) where it may also be possible.